![]() To that end, Mall and his colleagues not only prepared and preserved serial sections of the embryos, they also made hundreds of three-dimensional models at different stages of growth. He had more success basing his "staging" scheme on morphological characteristics. Mall took it upon himself to find a better way. It is very difficult to accurately age an embryo, and it could shrink a full 50% in the preserving fluids. Surprizingly age and size proves a poor way to organize embryos. (More? Franklin Mall)įranklin Mall Links: Franklin Mall | 1891 26 Day Human Embryo | 1905 Blood-Vessels of the Brain | 1906 Human Ossification | 1910 Manual of Human Embryology 1 | 1912 Manual of Human Embryology 2 | 1911 Mall Human Embryo Collection | 1912 Heart Development | 1915 Tubal Pregnancy | 1916 Human Magma in Normal and Pathological Development | 1917 Frequency Human Abnormalities | 1917 Human Embryo Cyclopia | 1918 Embryo Age | 1918 Appreciation | 1934 Franklin Mall biography PDF | Mall photograph | Mall painting | Mall painting | Carnegie Stages | Carnegie Embryos | Carnegie Collection | Category:Franklin Mall Mall began collecting human embryos while a postgraduate student in Lepzig with Wilhelm His, but didn't receive the first Carnegie specimen until his position at Johns Hopkins University. Mall (1862-1917) is most remembered for his work done at the Department of Embryology at the Carnegie Institute of Washington. The following text and information about the collection is modifed from the original Carnegie Institute website.įranklin P. To prevent confusion and consistency with the historic literature the Carnegie stages are used for human development on this site. There have been several other human systems of embryo categorisation developed, sometimes to establish a standard between species. Streeter also defined the fetal period as beginning when the humerus cartilage was replaced by bone marrow. ![]() Defined by the probability that more than 90 percent of the identifiable structures of the adult body have appeared by Carnegie stage 23. The division of human development into an embryonic (embryo) and fetal (fetus) periods was an historically based arbitrary system. Links: Embryonic Development | Carnegie Stage Comparison | Carnegie Institution - Contributions to Embryology | Human Embryo CollectionsĮmbryo Collections: Human Embryo Collections | Embryo Collections | Blechschmidt Collection | Carnegie Collection | Domenech-Mateu Collection | Harvard Collection | Hill Collection | Hinrichsen Collection | Hubrecht Collection | Kyoto Collection | Madrid Collection | Embryology Models | DEC Information | DECĬarnegie Stages: 1 | 2 | 3 | 4 | 5 | 6 | 7 | 8 | 9 | 10 | 11 | 12 | 13 | 14 | 15 | 16 | 17 | 18 | 19 | 20 | 21 | 22 | 23 | About Stages | Timeline Carnegie Embryos Scanning electron micrographs are published in collaboration with Prof Kathy Sulik. Note that many photographs of staged (using Carnegie criteria) human embryos on this current site are from the Kyoto collection in collaboration with Prof Kohei Shiota and Prof Shigehito Yamada. The table below also has detailed descriptions of each Carnegie stage as well as identifying embryo examples from different collections and the published literature. See also the timeline tabulation of both whole embryo and systematic development. Criteria beyond morphological features include ranges of age in days, number of somites present, and embryonic crown rump lengths (CRL). ![]() The human embryonic period proper is divided into 23 Carnegie stages covering the first 8 weeks post-ovulation ( GA week 10). ![]() Stages are based on the external and/or internal morphological development of the embryo, and are not directly dependent on either age or size. Carnegie stages are named after the famous US Institute which began collecting and classifying embryos in the early 1900's. ![]()
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